Gouania lupuloides

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Distribution Map: Based on vouchered plant specimens from wild populations. Cultivated occurrences are not mapped. View county names by placing the cursor over the map.

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Family:	RHAMNACEAE
Species:	Gouania lupuloides (L.)Urb.
Common Name:	CHEWSTICK; WHITEROOT
Status:	Native, FAC (NWPL)
Specimen:	View details of USF Herbarium specimens

** Not applicable or data not available.

Classification

Division
Class
Sub Class
Order	ROSALES
Family	RHAMNACEAE
Genus	Gouania
Species	Gouania lupuloides (L.)Urb. - CHEWSTICK; WHITEROOT

Citation

Citation	GOUANIA LUPULOIDES (Linnaeus) Urban, Symb. Antill. 4: 378. 1910.
Basionym:	Banisteria lupuloides Linnaeus 1753.
Type:	BARBADOS: Without data (lectotype: Plukenet, Phytographia t. 201(4). 1692). EPITYPE: BARBADOS: Kendal Gully, 24 Mar 1937, McIntosh 333 (epitype: K). Lecto- and epitypified by Pool, Ann. Missouri Bot. Gard 99: 520. 2014.

** Not applicable or data not available.

Source

County	Herbaria	Literature Citation (If Applicable)
Brevard	USF
Clay	FLAS
Indian River	USF
Manatee	USF
Martin	USF
Miami-Dade	USF
Monroe Keys	USF
Monroe Mainland	FNPS

Synonyms

– Denotes synonyms that are applicable to the state. Show these synonyms only

Synonym	Full Citation	Basionym	Type
Banisteria lupuloides	Banisteria lupuloides Linnaeus, Sp. Pl. 427. 1753.		TYPE: BARBADOS: Without data (lectotype: Plukenet, Phytographia t. 201(4). 1692). EPITYPE: BARBADOS: Kendal Gully, 24 Mar 1937, McIntosh 333 (epitype: K). Lecto- and epitypified by Pool, Ann. Missouri Bot. Gard 99: 520. 2014.
Gouania domingensis	Gouania domingensis Linnaeus, Sp. Pl., ed. 2. 1663. 1763, nom. illegit.	BASIONYM: Banisteria lupuloides Linnaeus 1753.
Gouania glabra	Gouania glabra Jacquin, Select. Stirp. Amer. Hist. 264. 1763, nom. illegit.	BASIONYM: Rhamnus domingensis Jacquin 1760.
Gouania glabriuscula	Gouania glabriuscula Stokes, Bot. Mat. Med. 1: 436. 1812, nom. Illegit.	BASIONYM: Banisteria lupuloides Linnaeus 1753.
Lupulus lupuloides	Lupulus lupuloides (Linnaeus) Kuntze, Revis. Gen. Pl. 1: 119. 1891.	BASIONYM: Banisteria lupuloides Linnaeus 1753.
Lupulus lupuloides var. domingensis	Lupulus lupuloides (Linnaeus) Kuntze, var. domingensis (Jacquin) Kuntze, Revis. Gen. Pl. 1: 119. 1891.	BASIONYM: Rhamnus domingensis Jacquin 1760.
Rhamnus domingensis	Rhamnus domingensis Jacquin, Enum. Syst. Pl. 17. 1760.		TYPE: Without data (neotype: Jacquin, Select. Stirp. Amer. Hist. t. 179(40). 1763). Neotypified by Pool, Ann. Missouri Bot. Gard. 99: 520. 2014.
Gouania cyclocarpa	Gouania cyclocarpa Smith, in Rees, Cycl. 16(2): Gouanai no. 3. 1811 ("1819").
Gouania lupuloides var. aptera	Gouania lupuloides (Linnaeus) Urban, var. aptera Urban, Symb. Antill. 4: 378. 1910.		TYPE: PUERTO RICO: Between Coama and Aibonito, 4 Nov 1885, Sintenis 1960 (lectotype: US; isolectotypes: BM, GH, K). Lectotypified by Pool, Ann. Missouri Bot. Gard. 99: 520. 2014.
Gouania lupuloides var. parvifolia	Gouania lupuloides (Linnaeus) Urban, var. parvifolia Hadac, Folia Geobot. Phytotax. 5: 436. 1970.		TYPE: CUBA: Las Villas: Playa larga, 0.5 km ad boreum a balneis, 22 Dec 1967, Hadac 838 (holotype: PR).
Gouania pubescens var. martinicensis	Gouania pubescens Poiret, var. martinicensis Poiret, in Lamarck, Encycl., Suppl. 2: 820. 1812 ("1811").		TYPE: MARTINIQUE: Without data (lectotype: Lamarck, tgabl. Encycl. T. 845 (1(h, i)). 1799 ("1793")). Lectotypified by Pool, Ann. Missouri Bot. Gard. 99: 520. 2014.
Gouania viridis	Gouania viridis Brandegee, Univ> Calif. Publ. Bot. 6: 501. 1919.		TYPE: MEXICO: Veracruz: Zacuapán, Oct 1917, Purpus 8032 (holtype: UC; isotypes: GH, MO, NY, US).
Lupulus lupuloides forma heterocarpus	Lupulus lupuloides (Linnaeus) Kuntze, forma heterocarpus Kuntze, Revis. Gen. Pl. 1: 119. 1891.		TYPE: COSTA RICA:

Specimens

Country	State	County	Date	Specimen	Notes
USA	Florida	Monroe Co.	13 Oct 1962	G. R. Cooley 9239
USA	Florida	Martin Co.	25 Jul 1990	R. Mears s.n.
USA	Florida	Miami-Dade Co.	19 Dec 1966	O. K. Lakela 30422
USA	Florida	Monroe Co.	12 Jan 1980	W. C. Brumbach 9537
USA	Florida	Monroe Co.	18 Jan 1969	R. W. Long 2785
USA	Florida	Monroe Co.	18 Jan 1969	R. W. Long 31760
USA	Florida	Monroe Co.	06 Dec 2000	K. A. Bradley 1309
USA	Florida	Martin Co.	Apr 1989	R. O. Woodbury s.n.
USA	Florida	Miami-Dade Co.	15 Oct 1962	G. R. Cooley 9356
USA	Florida	Indian River Co.	26 Nov 1978	R. P. Wunderlin 6483
USA	Florida	Monroe Co.	03 Oct 1962	F. C. Craighead s.n.
USA	Florida	Brevard Co.	s.d.	A. H. Curtiss 474
USA	Florida	Monroe Co.	15 Nov 1979	W. C. Brumbach 9528
USA	Florida	Monroe Co.	29 Apr 1963	F. C. Craighead s.n.
USA	Florida	Monroe Co.	27 Nov 1969	R. W. Long 2980
Jamaica	Middlesex Co.	St. Ann Par.	20-31 Dec 1953	R. A. Howard 13409
Jamaica	Cornwall Co.	St. Elizabeth Par.	Jun 1847	J. Wolle s.n.
Jamaica	Middlesex Co.	St. Catherine Par.	15 Nov 1953	G. R. Proctor 8188
USA	Florida	Brevard Co.	09 Dec 1929	M. Mulvania s.n.
Bahamas	Eleuthera Island		22 Dec 1969	W. H. Lewis 7431
Dominican Republic	La Altagracia Prov.		05 Feb 1981	T. A. Zanoni 10737	"Bejuco de Indio" or "Palo de Jabón"; common names here.
Puerto Rico		Villalba Mun.	28 Mar 1973	D. Burch s.n.
U.S. Virgin Islands	St. John Island		25 Aug 1994	J. E. Poppleton s.n.
Dominican Republic	Samana Prov.		07 Dec 1982	T. A. Zanoni 24703
Bahamas	Great Abaco Island		14 Dec 1979	R. P. Wunderlin 8309
USA	Florida	Miami-Dade Co.	15 Aug 2002	J. N. Burch 2605
Mexico	Yucatán		06 Jan 1982	S. Darwin 2170
Honduras	Francisco Morazán Dept.		06 Apr 1985	Z. Nolasco 157
Mexico	Yucatán		08 Oct 1983	R. P. Sauleda 8465
Mexico	Yucatán		07 Oct 1983	R. P. Sauleda 8451
Guyana	Upper Takutu-Upper Essequibo		21 Aug 1995	S. R. Hill 27322
Colombia	Caquetá Dept.		10 Jan 1974	G. Davidse 5651A
Peru	Amazonas		05 May 1973	E. Ancuash 327	Local name: tagkinik.
Mexico	Yucatán		09 Oct 1979	R. J. Gustafson 1476
USA	Florida	Monroe Co.	04 Jan 1958	M. L. Gilbert 61

Listed Status: Florida

Listed Status:	(Florida):Source - Plants in the Preservation of Native Flora of Florida Act. Chapter 5B-40, Florida Administrative Code. 1998, amended.
Endangered:	A species of plants native to the state that are in imminent danger of extinction within the state, the survival of which is unlikely if the causes of a decline in the number of plants continue, and includes all species determined to be endangered or threatened pursuant to the Federal Endangered Species Act of 1973, as amended.
Threatened:	Listed as Threatened Plants in the Preservation of Native Flora of Florida Act. Defined as species of plants native to the state that are in rapid decline in the number of plants within the state, but which have not so decreased in such number as to cause them to be endangered.

Listed Status: US

Listed Status: US	(U.S.) Source - List of Endangered and Threatened Wildlife and Plants under the provisions of the Endangered Species Act of 1973, as amended.
Endangered:	Any species which is in danger of extinction throughout all or a significant portion of its range.
Threatened:	Any species which is likely to become an endangered species within the foreseeable future throughout all or a significant portion of its range.

WAP: Wetland AssessmentProcedure

Wetland Assessment Procedure (WAP): Source - Southwest Florida Water Management District, Wetland Assessment Procedure Instruction Manual for Isolated Wetlands (March 2005).

AD:	Adaptive Species. Plant species designated as FAC or Upland by DEP, but commonly seen in the transition zone in limited numbers.
D:	Deep Species. Plant species commonly found in the deep zone, and designated either FAC or OBL by DEP.
OD:	Outer Deep Species. Plant species commonly found in the outer deep zone, and designated either FACW or OBL by DEP.
T:	Transition Species. Plant species commonly found in the transition zone, and designated either FACW or OBL by DEP.
U:	Upland Species. Plant species that are not expected to be seen in wetlands.

ANY - WAP critieria is not taken into consideration
Yes - Show results with WAP designations
No - Show results without WAP designations

FISC: Florida Invasive Species Council

FISC:

Source - Florida Invasive Species Council's 2017 List of Florida's Most Invasive Species

Category I - Species that are invading and disrupting native plant communities in Florida. This definition does not rely on the economic severity or geographic range of the problem, but on the documented ecological damage caused.

Category II - Species that have shown a potential to disrupt native plant communities. These species may become ranked as Category I, but have not yet demonstrated disruption of natural Florida communities.

DEP: Department of Environmental Protection

Wetland Status, Department of Environmental Regulation (DEP): Source - Delineation of the Landward Extent of Wetlands and Surface Waters, Chapter 62-340, Florida Administrative Code. 1994.

OBL:	Obligate wetland. Occurs almost always under natural conditions in wetlands.
FACW:	Facultative Wetland. Usually occurs in wetlands, but occasionally found in non-wetlands
FAC:	Facultative. equally likely to occur in wetlands or non-wetlands.

NWPL: National Wetland Plant List

National Wetland Plant List (NWPL): Source - Lichvar, R.W., M. Butterwick, N.C. Melvin, and W.N. Kirchner. 2014. The National Wetland Plant List: 2014 Update of Wetland Ratings. Phytoneuron 2014-41: 1-42.

OBL:	Obligate Wetland. Occurs almost always (estimated probability 99%) under natural conditions in wetlands.
FACW:	Facultative Wetland. Usually occurs in wetlands (estimated probability 67%-99%), but occasionally found in non-wetlands.
FAC:	Facultative. Equally likely to occur in wetlands or non-wetlands (estimated probability 34%-66%).
FACU:	Facultative Upland. Usually occurs in non-wetlands (estimated probability 67%-99%), but occasionally found in wetlands (estimated probability 1%-33%).
UPL:	Obligate Upland. Occurs in wetlands in another region, but occurs almost always (estimated probability 99%) under natural conditions in non-wetlands in the regions specified.

Persisting

Identifying species that appear as waifs or only periodically appear in the flora for a few seasons.

Any - Persisting critieria is not taken into consideration
Yes - Show results with a Persisting value of Y
No - Show results without Persisting value of N

Duration

Annual:	Plants that perform their entire lifecycle within a single growing season. All roots, stems, and leaves die at the end of the growing season. Over wintering seeds allow the next generation to appear.
Biennial:	A plant that is typically vegetative its first year and blooms the following season. Once it has bloomed and set seed, the plant dies. Garlic mustard is an example of a biennial.
Perennial:	These plants live for three or more seasons. Many perennials may not be mature enough to bloom during its first year. Perennial wildflowers re-grow each season from overwinter root material.

Category

Vascular:	Any of various plants that have the vascular tissues xylem and phloem. The vascular plants include all seed-bearing plants (the gymnosperms and angiosperms) and the pteridophytes (including the ferns, lycophytes, and horsetails). Also called tracheophyte.
Bryophyte:	A large group of seedless green plants including the mosses, liverworts, and hornworts. Bryophytes lack the specialized tissues xylem and phloem that circulate water and dissolved nutrients in the vascular plants. Bryophytes generally live on land but are mostly found in moist environments, for they have free-swimming sperm that require water for transport. In contrast to the vascular plants, the gametophyte (haploid) generation of bryophytes constitutes the larger plant form, while the small sporophyte (diploid) generation grows on or within the gametophyte and depends upon it for nutrition.
Lichen:	The mutualistic symbiotic association of a fungus with an alga or a cyanobacterium, or both. The fungal component of a lichen absorbs water and nutrients from the surroundings and provides a suitable environment for the alga or cyanobacterium. These live protected among the dense fungal hyphae and produce carbohydrates for the fungus by photosynthesis. Owing to this partnership, lichens can thrive in harsh environments such as mountaintops and polar regions. The more familiar lichens grow slowly as crusty patches, but lichens are found in a variety of forms, such as the tall, plantlike reindeer moss. The association between the different organisms in a lichen is so close that lichens are routinely referred to as a single organism, and scientists classify lichens using the name of the fungal component.

(Definitions from: American Heritage Science Dictionary)

Growth Habit

Intro paragraph to be provided by New York.

Vine:	DESCRIPTION TO BE PROVIDED BY NEW YORK.
Shrub:	DESCRIPTION TO BE PROVIDED BY NEW YORK.
Tree:	DESCRIPTION TO BE PROVIDED BY NEW YORK.
Et cetera:	DESCRIPTION TO BE PROVIDED BY NEW YORK.

State Rank

This numeric rank provides the relative rarity for each species based on a scale from 1 (very rare) to 5 (common). These ranks carry no legal status.

S1 - Typically 5 or fewer occurrences, very few remaining individuals, acres, or miles of stream, or some factor of its biology making it especially vulnerable in the state.
S2 - Typically 6 to 20 occurrences, few remaining individuals, acres, or miles of stream, or factors demonstrably making it very vulnerable in the state.
S3 - Typically 21 to 100 occurrences, limited acreage, or miles of stream in the state.
S4 - Apparently secure in the state.
S5 - Demonstrably secure in the state.
SE - State exotic or non-native.
SH - Historically known from the state, but not seen in the past 15 years.
SNA - Species for which a rank is not applicable. This is mainly those species which are now excluded from flora for various reasons.
SNR - Not yet ranked.
SX - Apparently extirpated from the state.

Global Rank

Each species' global rank is determined by NatureServe. These ranks carry no legal weight. The global rank reflects the species worldwide rarity.

G1 - Critically imperiled globally because of extreme rarity (5 or fewer occurrences), or very few remaining acres, or miles of stream) or especially vulnerable to extinction because of some factor of its biology.
G2 - Imperiled globally because of rarity (6 - 20 occurrences, or few remaining acres, or miles of stream) or very vulnerable to extinction throughout its range because of other factors.
G3 - Imperiled globally because of rarity (6 - 20 occurrences, or few remaining acres, or miles of stream) or very vulnerable to extinction throughout its range because of other factors.
G4 - Imperiled globally because of rarity (6 - 20 occurrences, or few remaining acres, or miles of stream) or very vulnerable to extinction throughout its range because of other factors.
G5 - Demonstrably secure globally, though it may be quite rare in parts of its range, especially at the periphery.
GH - Historically known, with the expectation that it might be rediscovered.
GNA - Species for which a rank is not applicable. NatureServe does not typically rank hybrid species.
GNR - Not yet ranked.
GX - Species believed to be extinct.

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Filter by County - Counties to Exclude - ANY versus ALL

Any - An Any search will combine the list of counties to exclude with a Boolean Or. Plant species returned will not be found within at least one of the selected counties.
All - An All search will combine the list of counties to exclude with a Boolean And. Plant species returned will not be found within any of the selected counties.

Associated Ecological Communities

View a List of All Ecological Communities

For more information, contact: Richard Wunderlin or Bruce Hansen, Alan Franck, or Karla Alvarado

Web Development: USF Water Institute

A member of the University of South Florida family of PlantAtlas.org websites

Citation Information:

Wunderlin, R. P., B. F. Hansen, A. R. Franck, and F. B. Essig. 2024. Atlas of Florida Plants (http://florida.plantatlas.usf.edu/). [S. M. Landry and K. N. Campbell (application development), USF Water Institute.] Institute for Systematic Botany, University of South Florida, Tampa.

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Definition: Vouchered Specimen

A voucher specimen is a pressed and thoroughly dried plant sample deposited in a herbarium, and is intended to be a permanent record supporting research purposes. A voucher may be a record of a plant's occurrence in a particular area, or a specific example of a plant used in a scientific investigation.

Proper vouchers display all the necessary attributes for complete identification of the plant, and are to be accompanied by accurate locality, habitat, collection time, and collector data.

Only plant populations vouchered by specimens deposited in Index Herbariorum http://sweetgum.nybg.org/science/ih/ recognized herbaria are represented on this map.