Cyperus articulatus

Distribution Map: Based on vouchered plant specimens from wild populations. Cultivated occurrences are not mapped. View county names by placing the cursor over the map.

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Family:	CYPERACEAE
Species:	Cyperus articulatus L.
Common Name:	JOINTED FLATSEDGE
Status:	Native, OBL (DEP), OBL (NWPL)
Specimen:	View details of USF Herbarium specimens

** Not applicable or data not available.

Classification

Division
Class
Sub Class
Order	POALES
Family	CYPERACEAE
Genus	Cyperus
Species	Cyperus articulatus L. - JOINTED FLATSEDGE

Citation

Citation	CYPERUS ARTICULATUS Linnaeus, Sp. Pl. 44. 1753.
Basionym:	**
Type:	JAMAICA: Without data (lectotype: Sloane, Voy. Jamaica 1: t. 81(1). 1707). Lectotypified by G. C. Tucker, Syst. Bot. Monogr. 2: 42. 1983.

** Not applicable or data not available.

Source

County	Herbaria	Literature Citation (If Applicable)
Brevard	USF
Charlotte	FSU
Citrus	USF
Clay	FLAS
Collier	USF
DeSoto	USF
Dixie	FSU
Duval	FLAS
Franklin	FSU
Hernando	USF
Highlands	USF
Hillsborough	USF
Lake	FLAS
Lee	USF
Manatee	USF
Marion	FLAS
Martin	USF
Miami-Dade	USF
Monroe Mainland	FNPS
Okeechobee	USF
Orange	USF
Osceola	FLAS
Palm Beach	USF
Polk	USF
Putnam	USF
Sarasota	FSU
St. Lucie	USF
Volusia	FSU
Wakulla	FSU

Synonyms

– Denotes synonyms that are applicable to the state. Show these synonyms only

Synonym	Full Citation	Basionym	Type
Chlorocyperus articulatus	Chlorocyperus articulatus (Linnaeus) Rikli, Jahrb. Wiss. Bot. 27: 563. 1895.	BASIONYM: Cyperus articulatus Linnaeus 1753.
Cyperus articulatus var. multiflorus	Cyperus articulatus Linnaeus, var. multiflorus Kükenthal, in Engler, Pflanzenr. 4(Heft 101): 79. 1935.		TYPE: FLORIDA: Lake Co.: Near Eustis, s.d., Nash 1701 (holotype: B?; isotypes: CONN, MICH, NY).
Chlorocyperus cordobensis	Chlorocyperus cordobensis Palla, Allg. Bot. Z. 9: 69. 1903.		TYPE: ARGENTINA:
Cyperus articulatus forma longispiculosus	Cyperus articulatus Linnaeus, forma longispiculosus Kükenthal, Repert. Spec. Nov. Regni Veg. 26: 250. 1929.
Cyperus articulatus var. conglomeratus	Cyperus articulatus Linnaeus, var. conglomeratus Britton, Bull. Torrey Bot. Club 13: 210. 1886.		TYPE: TEXAS: Valley of the lower Rio Grande, 1879-1883, Buckley s.n. (holortype: NY).
Cyperus articulatus var. erythrostachys	Cyperus articulatus Linnaeus, var. erythrostachys Graebner, Repert. Spec. Nov. Regni Veg. 16: 25. 1919.
Cyperus articulatus var. fistulosus	Cyperus articulatus Linnaeus, var. fistulosus Kükenthal, in Engler, Pflanzenr. 4(Heft 101): 79. 1935.
Cyperus articulatus var. nodosus	Cyperus articulatus Linnaeus, var. nodosus (Humboldt & Bonpland ex Willdenow) Kükenthal, in Engler, Pflanzenr. 4(Heft 101): 79. 1936.	BASIONYM: Cyperus nodosus Willdenow 1809.
Cyperus borbonicus	Cyperus borbonicus Steudel, Syn. Pl. Glumac. 2: 31. 1854.		TYPE: REUNION ISLAND:
Cyperus cordobensis	Cyperus cordobensis (Palla) Hicken, Darwiniana 1: 112. 1924.	BASIONYM: Chlorocyperus cordobensis Palla 1903.
Cyperus corymbosus	Cyperus corymbosus Rottbøll, Descr. Pl. Rar. 19. 1772.		TYPE: SURINAM: Without data, Rolander s.n. (holotype: C).
Cyperus corymbosus var. brevispiculatus	Cyperus corymbosus Rottbøll, var. brevispiculatus (Kuntze) Kükenthal, in Engler, Pflanzenr. 4(Heft 101): 82. 1936.	BASIONYM: Cyperus enodis Boeckeler, var. brevispiculatus Kuntze 1891.
Cyperus corymbosus var. longispiculatus	Cyperus corymbosus Rottbøll, var. longispiculatus (Kuntze) Kükenthal, in Engler, Pflanzenr. 4(Heft 101): 82. 1936.	BASIONYM: Cyperus enodis Boeckeler, var. longispiculatus Kuntze 1891.
Cyperus corymbosus var. subnodosus	Cyperus corymbosus Rottbøll, var. subnodosus (Nees von Esenbeck & Meyen) Kükenthal, Repert. Spec. Nov. Regni Veg. 23: 185. 1926.	BASIONYM: Cyperus subnodosus Nees von Esenbeck & Meyen 1842.
Cyperus diphyllus	Cyperus diphyllus Retzius, Observ. Bot. 8: 11. 1788.
Cyperus diphyllus var. elatior	Cyperus diphyllus Retzius, var. elatior Bentham, Fl. Austral. 7: 279. 1878.		TYPE: AUSTRALIA: Queensland:
Cyperus diphyllus var. triangularis	Cyperus diphyllus Retzius, var. triangularis Boeckeler, Linnaea 36: 273. 1870, nom. inadmiss.	BASIONYM: Cyperus diphyllus Retzius 1788.
Cyperus enodis	Cyperus enodis Boeckeler, Linnaea 36: 271. 1870.		TYPE: BURMA:
Cyperus enodis forma ferrugineus	Cyperus enodis Boeckeler, forma ferrugineus Kuntze, Revis. Gen. Pl. 2: 749. 1891.
Cyperus enodis forma fuscus	Cyperus enodis Boeckeler, forma fuscus Kuntze, Revis. Gen. Pl. 2: 749. 1891.
Cyperus enodis forma griseus	Cyperus enodis Boeckeler, forma griseus Kuntze, Revis. Gen. Pl. 2: 749. 1891.
Cyperus enodis var. brevispiculatus	Cyperus enodis Boeckeler, var. brevispiculatus Kuntze, Revis. Gen. Pl. 2: 749. 1891.
Cyperus enodis var. longispiculatus	Cyperus enodis Boeckeler, var. longispiculatus Kuntze, Revis. Gen. Pl. 2: 749. 1891.
Cyperus gula-methhi	Cyperus gula-methhi Schultes, Mant. 2: 125. 1824.	BASIONYM: Cyperus nudus Roxburgh 1820, non Kunth 1816.
Cyperus gymnos	Cyperus gymnos Schultes, Mant. 2: 97. 1824.
Cyperus interceptus	Cyperus interceptus Steudel, Syn. Pl. Glumac. 2: 31. 1854.
Cyperus koenigii	Cyperus koenigii Vahl, Enum. Pl. 2: 302. 1805.
Cyperus longus var. pallescens	Cyperus longus Linnaeus, var. pallescens (Desfontaines) Cosson & Durieu de Maisonneuve, Expl. Sci. Algérie, Bot. II. 2249. 1855.	BASIONYM: Cyperus pallescens Desfontaines 1798.
Cyperus niloticus	Cyperus niloticus Forsskål, Fl. Aegypt.-Arab. 13. 1775.		TYPE: EGYPT:
Cyperus nodosus	Cyperus nodosus Humboldt & Bonpland ex Willdenow, Enum. Pl. 72. 1809.		TYPE: PERU:
Cyperus nodosus var. aphyllus	Cyperus nodosus Willdenow, var. aphyllus Boeckeler, Linnaea 36: 275. 1870, nom. inadmiss.	BASIONYM: Cyperus nodosus Willdenow 1809.
Cyperus nodosus var. subnodosus	Cyperus nodosus Willdenow, var. subnososus (Nees von Esenbeck & Meyen) Boeckeler, Linnaea 36: 276. 1870.	BASIONYM: Cyperus subnodosus Nees von Esenbeck & Meyen 1842.
Cyperus nudus	Cyperus nudus Roxburgh, Fl. Ind. 1: 191. 1820, non Kunth 1816, nec Roxburgh p. 212, 1820.		TYPE: INDIA:
Cyperus nudus	Cyperus nudus Roxburgh, Fl. Ind., ed. 1832. 1: 209. 1832, non Kunth 1816, nec Roxburgh 1832.		TYPE: INDIA: Bengal:
Cyperus pallescens	Cyperus pallescens Desfontaines, Fl. Atlant. 1: 45, t. 9. 1798.
Cyperus roestelii	Cyperus roestelii Kunth, Enum. Pl. 2: 58. 1837.
Cyperus seminudus	Cyperus seminudus Roxburgh, Fl. Ind., ed. 1832. 1: 187. 1832.		TYPE: INDIA:
Cyperus subnodosus	Cyperus subnodosus Nees von Esenbeck & Meyen, Nov. Actorum Acad. Caes. Leop.-Carol. Nat. Cur. 19(Suppl. 1): 59. 1842.		TYPE: PERU:
Cyperus tagetiformis	Cyperus tagetiformis Roxburgh ex Arnott, in R. Wight, Contr. Bot. India 89. 1834.		TYPE: INDIA:
Cyperus tenuicomus	Cyperus tenuicomus Miquel, Linnaea 19: 223. 1846.

Specimens

Country	State	County	Date	Specimen
USA	Florida	Collier Co.	19 Jul 1965	O. K. Lakela 29000
USA	Florida	Collier Co.	06 Aug 1967	O. K. Lakela 30916
USA	Florida	Collier Co.	30 Jul 1968	O. K. Lakela 31530
USA	Florida	Martin Co.	Jul 1989	R. O. Woodbury s.n.
USA	Florida	DeSoto Co.	09 Oct 2011	R. Mears 5820
USA	Florida	Highlands Co.	06 Sep 1997	C. vanHoek 958
USA	Florida	Okeechobee Co.	22 Oct 1998	E. L. Bridges 25372
USA	Florida	Collier Co.	26 Jul 1978	J. Popenoe 1330
USA	Florida	Citrus Co.	24 Jun 1984	B. F. Hansen 9959
USA	Florida	Hillsborough Co.	10 Jul 1986	J. Massey s.n.
USA	Florida	Hillsborough Co.	13 Apr 1989	K. Holland s.n.
USA	Florida	Polk Co.	12 Jul 1979	A.G. Shuey 2248
USA	Florida	Okeechobee Co.	29 May 1980	B. F. Hansen 7300
USA	Florida	Citrus Co.	19 Jul 1958	R. Kral 7322
USA	Florida	Lee Co.	13 Sep 2007	J. M. Kunzer 2179
USA	Florida	Brevard Co.	19 Sep 1972	A.G. Shuey s.n.
USA	Florida	Palm Beach Co.	15 Jun 2009	R. Mears s.n.
USA	Florida	Highlands Co.	28 Oct 1997	S. L. Orzell 25255
USA	Florida	Orange Co.	10 Sep 1971	D. Auth s.n.
USA	Florida	Hillsborough Co.	13 Sep 1973	R. Wilkins s.n.
USA	Florida	Manatee Co.	06 Jun 1984	G. Robinson 643
USA	Florida	Hernando Co.	16 Jul 1989	R. Mears s.n.
USA	Florida	Brevard Co.	28 Jun 1973	A.G. Shuey s.n.
USA	Florida	Miami-Dade Co.	15 Jun 1958	W. G. Atwater GS-71
USA	Florida	St. Lucie Co.	05 Jul 2013	D. H. Williams 3714
USA	Louisiana	Jefferson Par.	26 Oct 1947	J. A. Ewan 17468
USA	Mississippi	Hancock Co.	08 Aug 1954	D. Demaree 36019
USA	Mississippi	Hancock Co.	10 Aug 1952	J. D. Ray, Jr. 3204
USA	Louisiana	Lafayette Par.	07 Sep 1979	K. A. Vincent 2862
USA	Florida	Hillsborough Co.	15 Oct 2009	S. Dickman 167
USA	Texas	Jefferson Co.	06 Aug 1983	R. D. Thomas 4537
Ecuador	Guayas Prov.		23 Jan 1983	C. H. Dodson 13631
Suriname	Paramaribo Distr.		26 Jun 1961	W. H. A. Hekking 879
Mexico	Veracruz	Alvarado Mun.	06 Jul 1980	B. F. Hansen 7653
USA	Florida	Putnam Co.	17 Jul 2017	E. A. Revuelta 129

Listed Status: Florida

Listed Status:	(Florida):Source - Plants in the Preservation of Native Flora of Florida Act. Chapter 5B-40, Florida Administrative Code. 1998, amended.
Endangered:	A species of plants native to the state that are in imminent danger of extinction within the state, the survival of which is unlikely if the causes of a decline in the number of plants continue, and includes all species determined to be endangered or threatened pursuant to the Federal Endangered Species Act of 1973, as amended.
Threatened:	Listed as Threatened Plants in the Preservation of Native Flora of Florida Act. Defined as species of plants native to the state that are in rapid decline in the number of plants within the state, but which have not so decreased in such number as to cause them to be endangered.

Listed Status: US

Listed Status: US	(U.S.) Source - List of Endangered and Threatened Wildlife and Plants under the provisions of the Endangered Species Act of 1973, as amended.
Endangered:	Any species which is in danger of extinction throughout all or a significant portion of its range.
Threatened:	Any species which is likely to become an endangered species within the foreseeable future throughout all or a significant portion of its range.

WAP: Wetland AssessmentProcedure

Wetland Assessment Procedure (WAP): Source - Southwest Florida Water Management District, Wetland Assessment Procedure Instruction Manual for Isolated Wetlands (March 2005).

AD:	Adaptive Species. Plant species designated as FAC or Upland by DEP, but commonly seen in the transition zone in limited numbers.
D:	Deep Species. Plant species commonly found in the deep zone, and designated either FAC or OBL by DEP.
OD:	Outer Deep Species. Plant species commonly found in the outer deep zone, and designated either FACW or OBL by DEP.
T:	Transition Species. Plant species commonly found in the transition zone, and designated either FACW or OBL by DEP.
U:	Upland Species. Plant species that are not expected to be seen in wetlands.

ANY - WAP critieria is not taken into consideration
Yes - Show results with WAP designations
No - Show results without WAP designations

FISC: Florida Invasive Species Council

FISC:

Source - Florida Invasive Species Council's 2017 List of Florida's Most Invasive Species

Category I - Species that are invading and disrupting native plant communities in Florida. This definition does not rely on the economic severity or geographic range of the problem, but on the documented ecological damage caused.

Category II - Species that have shown a potential to disrupt native plant communities. These species may become ranked as Category I, but have not yet demonstrated disruption of natural Florida communities.

DEP: Department of Environmental Protection

Wetland Status, Department of Environmental Regulation (DEP): Source - Delineation of the Landward Extent of Wetlands and Surface Waters, Chapter 62-340, Florida Administrative Code. 1994.

OBL:	Obligate wetland. Occurs almost always under natural conditions in wetlands.
FACW:	Facultative Wetland. Usually occurs in wetlands, but occasionally found in non-wetlands
FAC:	Facultative. equally likely to occur in wetlands or non-wetlands.

NWPL: National Wetland Plant List

National Wetland Plant List (NWPL): Source - Lichvar, R.W., M. Butterwick, N.C. Melvin, and W.N. Kirchner. 2014. The National Wetland Plant List: 2014 Update of Wetland Ratings. Phytoneuron 2014-41: 1-42.

OBL:	Obligate Wetland. Occurs almost always (estimated probability 99%) under natural conditions in wetlands.
FACW:	Facultative Wetland. Usually occurs in wetlands (estimated probability 67%-99%), but occasionally found in non-wetlands.
FAC:	Facultative. Equally likely to occur in wetlands or non-wetlands (estimated probability 34%-66%).
FACU:	Facultative Upland. Usually occurs in non-wetlands (estimated probability 67%-99%), but occasionally found in wetlands (estimated probability 1%-33%).
UPL:	Obligate Upland. Occurs in wetlands in another region, but occurs almost always (estimated probability 99%) under natural conditions in non-wetlands in the regions specified.

Persisting

Identifying species that appear as waifs or only periodically appear in the flora for a few seasons.

Any - Persisting critieria is not taken into consideration
Yes - Show results with a Persisting value of Y
No - Show results without Persisting value of N

Duration

Annual:	Plants that perform their entire lifecycle within a single growing season. All roots, stems, and leaves die at the end of the growing season. Over wintering seeds allow the next generation to appear.
Biennial:	A plant that is typically vegetative its first year and blooms the following season. Once it has bloomed and set seed, the plant dies. Garlic mustard is an example of a biennial.
Perennial:	These plants live for three or more seasons. Many perennials may not be mature enough to bloom during its first year. Perennial wildflowers re-grow each season from overwinter root material.

Category

Vascular:	Any of various plants that have the vascular tissues xylem and phloem. The vascular plants include all seed-bearing plants (the gymnosperms and angiosperms) and the pteridophytes (including the ferns, lycophytes, and horsetails). Also called tracheophyte.
Bryophyte:	A large group of seedless green plants including the mosses, liverworts, and hornworts. Bryophytes lack the specialized tissues xylem and phloem that circulate water and dissolved nutrients in the vascular plants. Bryophytes generally live on land but are mostly found in moist environments, for they have free-swimming sperm that require water for transport. In contrast to the vascular plants, the gametophyte (haploid) generation of bryophytes constitutes the larger plant form, while the small sporophyte (diploid) generation grows on or within the gametophyte and depends upon it for nutrition.
Lichen:	The mutualistic symbiotic association of a fungus with an alga or a cyanobacterium, or both. The fungal component of a lichen absorbs water and nutrients from the surroundings and provides a suitable environment for the alga or cyanobacterium. These live protected among the dense fungal hyphae and produce carbohydrates for the fungus by photosynthesis. Owing to this partnership, lichens can thrive in harsh environments such as mountaintops and polar regions. The more familiar lichens grow slowly as crusty patches, but lichens are found in a variety of forms, such as the tall, plantlike reindeer moss. The association between the different organisms in a lichen is so close that lichens are routinely referred to as a single organism, and scientists classify lichens using the name of the fungal component.

(Definitions from: American Heritage Science Dictionary)

Growth Habit

Intro paragraph to be provided by New York.

Vine:	DESCRIPTION TO BE PROVIDED BY NEW YORK.
Shrub:	DESCRIPTION TO BE PROVIDED BY NEW YORK.
Tree:	DESCRIPTION TO BE PROVIDED BY NEW YORK.
Et cetera:	DESCRIPTION TO BE PROVIDED BY NEW YORK.

State Rank

This numeric rank provides the relative rarity for each species based on a scale from 1 (very rare) to 5 (common). These ranks carry no legal status.

S1 - Typically 5 or fewer occurrences, very few remaining individuals, acres, or miles of stream, or some factor of its biology making it especially vulnerable in the state.
S2 - Typically 6 to 20 occurrences, few remaining individuals, acres, or miles of stream, or factors demonstrably making it very vulnerable in the state.
S3 - Typically 21 to 100 occurrences, limited acreage, or miles of stream in the state.
S4 - Apparently secure in the state.
S5 - Demonstrably secure in the state.
SE - State exotic or non-native.
SH - Historically known from the state, but not seen in the past 15 years.
SNA - Species for which a rank is not applicable. This is mainly those species which are now excluded from flora for various reasons.
SNR - Not yet ranked.
SX - Apparently extirpated from the state.

Global Rank

Each species' global rank is determined by NatureServe. These ranks carry no legal weight. The global rank reflects the species worldwide rarity.

G1 - Critically imperiled globally because of extreme rarity (5 or fewer occurrences), or very few remaining acres, or miles of stream) or especially vulnerable to extinction because of some factor of its biology.
G2 - Imperiled globally because of rarity (6 - 20 occurrences, or few remaining acres, or miles of stream) or very vulnerable to extinction throughout its range because of other factors.
G3 - Imperiled globally because of rarity (6 - 20 occurrences, or few remaining acres, or miles of stream) or very vulnerable to extinction throughout its range because of other factors.
G4 - Imperiled globally because of rarity (6 - 20 occurrences, or few remaining acres, or miles of stream) or very vulnerable to extinction throughout its range because of other factors.
G5 - Demonstrably secure globally, though it may be quite rare in parts of its range, especially at the periphery.
GH - Historically known, with the expectation that it might be rediscovered.
GNA - Species for which a rank is not applicable. NatureServe does not typically rank hybrid species.
GNR - Not yet ranked.
GX - Species believed to be extinct.

Filter by County - Counties to Include - ANY versus ALL

Any - An Any search will combine the list of counties to include with a Boolean Or. Plant species returned will be found within at least one of the selected counties.
All - An All search will combine the list of counties to include with a Boolean And. Plant species returned will be found within each of the selected counties.

Filter by County - Counties to Exclude - ANY versus ALL

Any - An Any search will combine the list of counties to exclude with a Boolean Or. Plant species returned will not be found within at least one of the selected counties.
All - An All search will combine the list of counties to exclude with a Boolean And. Plant species returned will not be found within any of the selected counties.

Associated Ecological Communities

View a List of All Ecological Communities

For more information, contact: Richard Wunderlin or Bruce Hansen, Alan Franck, or Karla Alvarado

Web Development: USF Water Institute

A member of the University of South Florida family of PlantAtlas.org websites

Citation Information:

Wunderlin, R. P., B. F. Hansen, A. R. Franck, and F. B. Essig. 2024. Atlas of Florida Plants (http://florida.plantatlas.usf.edu/). [S. M. Landry and K. N. Campbell (application development), USF Water Institute.] Institute for Systematic Botany, University of South Florida, Tampa.

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Definition: Vouchered Specimen

A voucher specimen is a pressed and thoroughly dried plant sample deposited in a herbarium, and is intended to be a permanent record supporting research purposes. A voucher may be a record of a plant's occurrence in a particular area, or a specific example of a plant used in a scientific investigation.

Proper vouchers display all the necessary attributes for complete identification of the plant, and are to be accompanied by accurate locality, habitat, collection time, and collector data.

Only plant populations vouchered by specimens deposited in Index Herbariorum http://sweetgum.nybg.org/science/ih/ recognized herbaria are represented on this map.