Thelypteris resinifera

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Family:	THELYPTERIDACEAE
Species:	Thelypteris resinifera (Desv.)Proctor
Common Name:	GLANDULAR MAIDEN FERN
Plant Notes:	Recent taxonomies place this species in the genus Amauropelta (Alvarez-Fuentes 2010; Almeida et al. 2016; PPG 2016).
Status:	Native, FACW (DEP), FACW (NWPL)
Specimen:	View details of USF Herbarium specimens

** Not applicable or data not available.

Classification

Division
Class
Sub Class
Order	POLYPODIALES
Family	THELYPTERIDACEAE
Genus	Thelypteris
Species	Thelypteris resinifera (Desv.)Proctor - GLANDULAR MAIDEN FERN

Citation

Citation	THELYPTERIS RESINIFERA (Desvaux) Proctor, Bull. Inst. Jamaica, Sci. Ser. 5: 63. 1953.
Basionym:	Polypodium resiniferum Desvaux 1811.
Type:	Without data (holotype: P).

** Not applicable or data not available.

Source

County	Herbaria	Literature Citation (If Applicable)
DeSoto	USF
Hillsborough	USF
Pasco	FLAS
Polk	USF

Synonyms

– Denotes synonyms that are applicable to the state. Show these synonyms only

Synonym	Full Citation	Basionym	Type
Amauropelta resinifera	Amauropelta resinifera (Desvaux) Pichi Sermolli, Webbia 31(1): 251. 1977.	BASIONYM: Polypodium reiniferum Desvaux 1811.
Aspidium conterminum var. strigosum	Aspidium conterminum Willdenow, var. strigosum D. C. Eaton, Bull. Torrey Bot. Club 7: 62. 1880.	BASIONYM: Aspidium strigosum Fée 1866, non Willdenow 1810, nec Mettenius 1858.
Aspidium strigosum	Aspidium strigosum Fée, Mém. Foug. 11: 78, pl. 22. 1866, non Willdenow 1810, nec Mettenius 1858.
Dryopteris contermina var. strigosa	Dryopteris contermina (Willdenow) Kuntze, var. strigosa (D. C. Eaton) Underwood, Native Ferns, ed. 4. 113. 1893.	BASIONYM: Aspidium conterminum Willdenow, var. strigosa D. C. Eaton 1880; Aspidium strigosum Fée 1866, non Willdenow 1810, nec Mettenius 1858.
Dryopteris panamensis	Dryopteris panamensis (C. Presl) C. Christensen, Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 7. 4: 292. 1907.	BASIONYM: Nephrodium panamense C. Presl 1825.
Dryopteris resinifera	Dryopteris resinifera (Desvaux) Weatherby, Contr. Gray Herb. 114. 32. 1936.	BASIONYM: Polypodium resiniferum Desvaux 1811.
Filix opposita var. strigosa	Filix opposita (Vahl) Farwell, var. strigosa (D. C. Eaton) Farwell, Rep. (Annual) Michigan Acad. Sci. 18: 82. 1916.	BASIONYM: Aspidium conterminum Willdenow, var. strigosa D. C. Eaton 1880; Aspidium strigosum Fée 1866, non Willdenow 1810, nec Mettenius 1858.
Lastrea resinifera	Lastrea resinifera (Desvaux) Copeland, Gen. Filic. 140. 1947.	BASIONYM: Polypodium resiniferum Desvaux 1811.
Nephrodium conterminum var. strigosum	Nephrodium conterminum (Willdenow) Desvaux, var. strigosum (D. C. Eaton) Gilbert, List N. Amer. Pterid. 17. 1901.	BASIONYM: Aspidium conterminum Willdenow, var. strigosa D. C. Eaton 1880; Aspidium strigosum Fée 1866, non Willdenow 1810, nec Mettenius 1858.
Nephrodium panamense	Nephrodium panamense C. Presl, Reliq. Haenk. 1: 35. 1825.		TYPE: PANAMA: Without data, Haenke s.n. (holotype: PR?; isotype: K?).
Polypodium litigiosum	Polypodium litigiosum Liebmann, Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 5. 1: 205. 1849.		TYPE: MEXICO: Veracruz: Mirador, s.d., Liebmann s.n. (holotype: C; isotype: K).
Polypodium resiniferum	Polypodium resiniferum Desvaux, Ges. Naturf. Freunde Berlin Mag. Neuesten Entdeck. Geseammten Naturk. 5: 317. 1810.		TYPE: Without data (holotype: P).
Thelypteris panamensis	Thelypteris panamensis (C. Presl) E. P. St. John, Amer. Fern J. 26: 44. 1936.	BASIONYM: Nephrodium panamense C. Presl 1825.
Aspidium exsudans	Aspidium exsudans E. Fournier, Mexic. Pl. 1: 93. 1872.		TYPE: MEXICO:
Aspidium exsudans var. feei	Aspidium exsudans E. Fournier, var. feei E. Fournier, Mexic. Pl. 1: 93. 1872.		TYPE: MEXICO:
Aspidium exsudans var. majus	Aspidium exsudans E. Fournier, var. majus E. Fournier, Mexic. Pl. 1: 93. 1872.		TYPE: MEXICO:
Aspidium ghiesbreghtii	Aspidium ghiesbreghtii Fournier, Mexic. Pl. 1: 94. 1872.		TYPE: MEXICO:
Dryopteris aequatorialis	Dryopteris aequatorialis Copeland, Univ. Calif. Publ. Bot. 19: 298, t. 53. 1941.		TYPE: ECUADOR: Esmeraldas: Parroquia de Concepción, below Playa Rica, 23 Dec 1936, Mexia 8486 (holotype: UC; isotypes: F, GH, MO, S, US).
Dryopteris litigiosa	Dryopteris litigiosa (Liebmann) C. Christensen ex H. Christ, Bull. Herb. Boissier, ser. 2. 7: 263. 1907.	BASIONYM: Polypodium litigiosum Liebmann 1849.
Dryopteris panamensis var. gonzalezii	Dryopteris panamensis (C. Presl) C. Christensen, var. gonzalezii C. Christensen, Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. math. Afd., ser. 7. 4: 294. 1907.		TYPE: MEXICO: Oaxaca: Cerro de San Felikpe, Oct 1900, González s.n. (holotype: P).
Dryopteris pseudosancta	Dryopteris pseudosancta C. Christensen, Smithsonian Misc. Collect. 52: 378. 1909.		TYPE: COSTA RICA: Río Toro Amarillo, Llanuras de Santa Clara, Apr 1896, Donnell Smith 6902 (holotype: US).
Lastrea aequatorialis	Lastrea aequatorialis (Copeland) Copeland, Gen. Fil. 138. 1947.	BASIONYM: Dryopteris aequatorialis Copeland 1941.
Lastrea leiboldiana	Lastrea leiboldiana C. Presl, Epimel. Bot. 41. 1851 ("1849").		TYPE: MEXICO:
Thelypteris aequatorialis	Thelypteris aequatorialis (Copeland) C. F. Reed, Phytologia 17: 258. 1968.	BASIONYM: Dryopteris aequatorialis Copeland 1941.
Thelypteris pseudosancta	Thelypteris pseudosancta (C. Christensen) C. F. Reed, Phytologia 17: 306. 1968.	BASIONYM: Dryopteris pseudosancta C. Christensen 1909.

Specimens

Country	State	County	Date	Specimen	Notes
USA	Florida	Hillsborough Co.	07 Dec 1953	R. Garrett s.n.
USA	Florida	Polk Co.	1933	Mrs. Peterson s.n.
USA	Florida	Polk Co.	02 May 1942	W. M. Buswell s.n.
USA	Florida	Polk Co.	09 Apr 1939	W. S. Phillips s.n.
USA	Florida	Polk Co.	17 Dec 1931	O. E. Jennings s.n.
USA	Florida	Polk Co.	24 Dec 1929	O. E. Jennings s.n.
USA	Florida	DeSoto Co.	17 Oct 1974	A.G. Shuey s.n.
USA	Florida	Polk Co.	19 Jun 1970	J. H. Scudder 145
USA	Florida	Polk Co.	18 Dec 1975	J. H. Scudder 253
USA	Florida	Polk Co.	19 Jun 1970	J. H. Scudder 146-A
USA	Florida	Hillsborough Co.	05 Jun 1980	A.G. Shuey 2395
USA	Florida	Hillsborough Co.	29 Jun 2015	S. Dickman s.n.
Jamaica	Middlesex Co.	Clarendon Par.	10 Apr 1953	G. R. Proctor 7843
Jamaica	Surrey Co.	St. Andrew Par.	10 May 1950	G. R. Proctor 4428	Cooley 946.
Jamaica	Surrey Co.	Portland Par.	20 Jul 1952	G. R. Cooley 1561
Honduras	Francisco Morazán Dept.		13 May 1984	L. López 194
Mexico	Veracruz		28 Jun 1980	M. Nee 18633
Honduras	Francisco Morazán Dept.		20 May 1984	E. J. Paredes 176
Honduras	Francisco Morazán Dept.		13 May 1984	L. López 193
Ecuador	Los Ríos Prov.		28 Jul 1980	B. F. Hansen 7752
Honduras	Francisco Morazán Dept.		19 Mar 1984	N. Cristoff 128
Honduras	Olancho Dept.		04 May 1985	S. Solabarrieta 161
Costa Rica	Puntarenas Prov.		17 May 1971	D. Burch 4479
USA	Florida	Hillsborough Co.	19 Jan 2015	S. Dickman 907
USA	Florida	Hillsborough Co.	22 Jul 2015	S. Dickman 986

Listed Status: Florida

Listed Status:	(Florida):Source - Plants in the Preservation of Native Flora of Florida Act. Chapter 5B-40, Florida Administrative Code. 1998, amended.
Endangered:	A species of plants native to the state that are in imminent danger of extinction within the state, the survival of which is unlikely if the causes of a decline in the number of plants continue, and includes all species determined to be endangered or threatened pursuant to the Federal Endangered Species Act of 1973, as amended.
Threatened:	Listed as Threatened Plants in the Preservation of Native Flora of Florida Act. Defined as species of plants native to the state that are in rapid decline in the number of plants within the state, but which have not so decreased in such number as to cause them to be endangered.

Listed Status: US

Listed Status: US	(U.S.) Source - List of Endangered and Threatened Wildlife and Plants under the provisions of the Endangered Species Act of 1973, as amended.
Endangered:	Any species which is in danger of extinction throughout all or a significant portion of its range.
Threatened:	Any species which is likely to become an endangered species within the foreseeable future throughout all or a significant portion of its range.

WAP: Wetland AssessmentProcedure

Wetland Assessment Procedure (WAP): Source - Southwest Florida Water Management District, Wetland Assessment Procedure Instruction Manual for Isolated Wetlands (March 2005).

AD:	Adaptive Species. Plant species designated as FAC or Upland by DEP, but commonly seen in the transition zone in limited numbers.
D:	Deep Species. Plant species commonly found in the deep zone, and designated either FAC or OBL by DEP.
OD:	Outer Deep Species. Plant species commonly found in the outer deep zone, and designated either FACW or OBL by DEP.
T:	Transition Species. Plant species commonly found in the transition zone, and designated either FACW or OBL by DEP.
U:	Upland Species. Plant species that are not expected to be seen in wetlands.

ANY - WAP critieria is not taken into consideration
Yes - Show results with WAP designations
No - Show results without WAP designations

FISC: Florida Invasive Species Council

FISC:

Source - Florida Invasive Species Council's 2017 List of Florida's Most Invasive Species

Category I - Species that are invading and disrupting native plant communities in Florida. This definition does not rely on the economic severity or geographic range of the problem, but on the documented ecological damage caused.

Category II - Species that have shown a potential to disrupt native plant communities. These species may become ranked as Category I, but have not yet demonstrated disruption of natural Florida communities.

DEP: Department of Environmental Protection

Wetland Status, Department of Environmental Regulation (DEP): Source - Delineation of the Landward Extent of Wetlands and Surface Waters, Chapter 62-340, Florida Administrative Code. 1994.

OBL:	Obligate wetland. Occurs almost always under natural conditions in wetlands.
FACW:	Facultative Wetland. Usually occurs in wetlands, but occasionally found in non-wetlands
FAC:	Facultative. equally likely to occur in wetlands or non-wetlands.

NWPL: National Wetland Plant List

National Wetland Plant List (NWPL): Source - Lichvar, R.W., M. Butterwick, N.C. Melvin, and W.N. Kirchner. 2014. The National Wetland Plant List: 2014 Update of Wetland Ratings. Phytoneuron 2014-41: 1-42.

OBL:	Obligate Wetland. Occurs almost always (estimated probability 99%) under natural conditions in wetlands.
FACW:	Facultative Wetland. Usually occurs in wetlands (estimated probability 67%-99%), but occasionally found in non-wetlands.
FAC:	Facultative. Equally likely to occur in wetlands or non-wetlands (estimated probability 34%-66%).
FACU:	Facultative Upland. Usually occurs in non-wetlands (estimated probability 67%-99%), but occasionally found in wetlands (estimated probability 1%-33%).
UPL:	Obligate Upland. Occurs in wetlands in another region, but occurs almost always (estimated probability 99%) under natural conditions in non-wetlands in the regions specified.

Persisting

Identifying species that appear as waifs or only periodically appear in the flora for a few seasons.

Any - Persisting critieria is not taken into consideration
Yes - Show results with a Persisting value of Y
No - Show results without Persisting value of N

Duration

Annual:	Plants that perform their entire lifecycle within a single growing season. All roots, stems, and leaves die at the end of the growing season. Over wintering seeds allow the next generation to appear.
Biennial:	A plant that is typically vegetative its first year and blooms the following season. Once it has bloomed and set seed, the plant dies. Garlic mustard is an example of a biennial.
Perennial:	These plants live for three or more seasons. Many perennials may not be mature enough to bloom during its first year. Perennial wildflowers re-grow each season from overwinter root material.

Category

Vascular:	Any of various plants that have the vascular tissues xylem and phloem. The vascular plants include all seed-bearing plants (the gymnosperms and angiosperms) and the pteridophytes (including the ferns, lycophytes, and horsetails). Also called tracheophyte.
Bryophyte:	A large group of seedless green plants including the mosses, liverworts, and hornworts. Bryophytes lack the specialized tissues xylem and phloem that circulate water and dissolved nutrients in the vascular plants. Bryophytes generally live on land but are mostly found in moist environments, for they have free-swimming sperm that require water for transport. In contrast to the vascular plants, the gametophyte (haploid) generation of bryophytes constitutes the larger plant form, while the small sporophyte (diploid) generation grows on or within the gametophyte and depends upon it for nutrition.
Lichen:	The mutualistic symbiotic association of a fungus with an alga or a cyanobacterium, or both. The fungal component of a lichen absorbs water and nutrients from the surroundings and provides a suitable environment for the alga or cyanobacterium. These live protected among the dense fungal hyphae and produce carbohydrates for the fungus by photosynthesis. Owing to this partnership, lichens can thrive in harsh environments such as mountaintops and polar regions. The more familiar lichens grow slowly as crusty patches, but lichens are found in a variety of forms, such as the tall, plantlike reindeer moss. The association between the different organisms in a lichen is so close that lichens are routinely referred to as a single organism, and scientists classify lichens using the name of the fungal component.

(Definitions from: American Heritage Science Dictionary)

Growth Habit

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Shrub:	DESCRIPTION TO BE PROVIDED BY NEW YORK.
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State Rank

This numeric rank provides the relative rarity for each species based on a scale from 1 (very rare) to 5 (common). These ranks carry no legal status.

S1 - Typically 5 or fewer occurrences, very few remaining individuals, acres, or miles of stream, or some factor of its biology making it especially vulnerable in the state.
S2 - Typically 6 to 20 occurrences, few remaining individuals, acres, or miles of stream, or factors demonstrably making it very vulnerable in the state.
S3 - Typically 21 to 100 occurrences, limited acreage, or miles of stream in the state.
S4 - Apparently secure in the state.
S5 - Demonstrably secure in the state.
SE - State exotic or non-native.
SH - Historically known from the state, but not seen in the past 15 years.
SNA - Species for which a rank is not applicable. This is mainly those species which are now excluded from flora for various reasons.
SNR - Not yet ranked.
SX - Apparently extirpated from the state.

Global Rank

Each species' global rank is determined by NatureServe. These ranks carry no legal weight. The global rank reflects the species worldwide rarity.

G1 - Critically imperiled globally because of extreme rarity (5 or fewer occurrences), or very few remaining acres, or miles of stream) or especially vulnerable to extinction because of some factor of its biology.
G2 - Imperiled globally because of rarity (6 - 20 occurrences, or few remaining acres, or miles of stream) or very vulnerable to extinction throughout its range because of other factors.
G3 - Imperiled globally because of rarity (6 - 20 occurrences, or few remaining acres, or miles of stream) or very vulnerable to extinction throughout its range because of other factors.
G4 - Imperiled globally because of rarity (6 - 20 occurrences, or few remaining acres, or miles of stream) or very vulnerable to extinction throughout its range because of other factors.
G5 - Demonstrably secure globally, though it may be quite rare in parts of its range, especially at the periphery.
GH - Historically known, with the expectation that it might be rediscovered.
GNA - Species for which a rank is not applicable. NatureServe does not typically rank hybrid species.
GNR - Not yet ranked.
GX - Species believed to be extinct.

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Associated Ecological Communities

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Citation Information:

Wunderlin, R. P., B. F. Hansen, A. R. Franck, and F. B. Essig. 2024. Atlas of Florida Plants (http://florida.plantatlas.usf.edu/). [S. M. Landry and K. N. Campbell (application development), USF Water Institute.] Institute for Systematic Botany, University of South Florida, Tampa.

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Definition: Vouchered Specimen

A voucher specimen is a pressed and thoroughly dried plant sample deposited in a herbarium, and is intended to be a permanent record supporting research purposes. A voucher may be a record of a plant's occurrence in a particular area, or a specific example of a plant used in a scientific investigation.

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