Najas marina

Distribution Map: Based on vouchered plant specimens from wild populations. Cultivated occurrences are not mapped. View county names by placing the cursor over the map.

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Family:	HYDROCHARITACEAE
Species:	Najas marina L.
Common Name:	SPINY WATERNYMPH
Status:	Native, OBL (NWPL)
Specimen:	View details of USF Herbarium specimens

** Not applicable or data not available.

Classification

Division
Class
Sub Class
Order	ALISMATALES
Family	HYDROCHARITACEAE
Genus	Najas
Species	Najas marina L. - SPINY WATERNYMPH

Citation

Citation	NAJAS MARINA Linnaeus, Sp. Pl. 1015. 1753.
Basionym:	**
Type:	"Europae maribus", without data (lectotype: LINN 1156.1). Lectotypified by Viinikka, Ann. Bot. Fenn. 13: 128. 1976.

** Not applicable or data not available.

Source

County	Herbaria	Literature Citation (If Applicable)
Brevard	USF
Collier	FLAS
Escambia	USF
Hillsborough	USF
Lee	USF
Levy	FLAS
Manatee	USF
Marion	FLAS
Miami-Dade	USF
Palm Beach	FLAS
Sarasota	USF
Wakulla	FTG

Synonyms

– Denotes synonyms that are applicable to the state. Show these synonyms only

Synonym	Full Citation	Basionym	Type
Ittnera najas	Ittnera najas C. C. Gmelin, Fl. Bad. 3: 590. 1808, nom. illegit.	BASIONYM: Najas marina Linnaeus 1753.
Najas gracilis	Najas gracilis (Morong) Small, Fl. S.E. U.S. 40, 1326. 1903.	BASIONYM: Najas major Allioni, var. gracilis Morong 1885.
Najas major	Najas major Allioni, Auct. Syn. Meth. Stirp. Horti Regii Taur. 3. 1773.		TYPE: Without data (lectotype: Micheli< Nov. Pl. Gen. t. 8(2). 1729). Lectotypified by Casper, Feddes Repert. 90: 217. 1979.
Najas major var. angustifolia	Najas major Allioni, var. angustifolia A. Braun, J. Bot. 2: 276. 1864.
Najas major var. gracilis	Najas major Allioni, var. gracilis Morong, Bot. Gaz. 10: 255. 1885.		TYPE: FLORIDA: Collier Co.: Palm Creek, near Cape Romano, Jul, Curtiss 2705 (holotype: NY; isotype: NY, NY).
Najas marina subsp. major	Najas marina Linnaeus, subsp. major (Allioni) Viinika, Ann. Bot. Fenn, 13: 129. 1976.	BASIONYM: Najas major Allioni 1773.
Najas marina var. angustifolia	Najas marina Linnaeus, var. angustifolia (A. Braun) K. Schumann, in Martius, Fl. Bras. 3(3): 725. 1894.	BASIONYM: Najas major Allioni, var. angustifolia A. Braun 1864.
Najas marina var. angustissima	Najas marina Linnaeus, var. angustissima K. Schumann, in Martius, Fl. Bras. 3(3): 725. 1894.		TYPE: FLORIDA: Collier Co.: "Palm Creek", near Cape Romano, s.d., Curtiss s.n. (holotype: ?).
Najas marina var. genuina	Najas marina Linnaeus, var. genuina K. Schumann, in Martius, Fl. Bras. 3(3): 724. 1894, nom. inadmiss.	BASIONYM: Najas marina Linnaeus 1753.
Najas marina var. gracilis	Najas marina Linnaeus, var. gracilis (Morong) Dudley, Cornell Univ. Sci. Bull. 2: 104. 1886.	BASIONYM: Najas major Allioni, var. gracilis Morong 1885.
Najas intermedia	Najas intermedia Gorski, Naturhist. Skizze 126. 1830.		TYPE: LITHUANIA: Lake Nawy near Wysoki-Dwor (10 mi. from Vilna), and around Strawa (4 mi. from Vilna), 1826, Gorski s.n. (lectotype: WI). Lectotypified by Bräuchler, Taxon 64: 1029. 2015.
Najas latifolia	Najas latifolia A. Braun, J. Bot. 2: 276. 1864.		TYPE: VENEZUELA:
Najas major subsp. intermedia	Najas major Allioni, subsp. intermedia (Gorski) K. Richter, Fl. Eur. 1: 18. 1890.	BASIONYM: Najas intermedia Gorski 1830.
Najas major subsp. multidentata	Najas major Allioni, subsp. multidentata (A. Braun) K. Richter, Fl. Eur. 1: 18. 1890.	BASIONYM: Najas major Allioni, var. multidentata A. Braun 1864.
Najas major subsp. paucidentata	Najas major Allioni, subsp. paucidentata (A. Braun) K. Richter, Fl. Eur. 1: 18. 1890.	BASIONYM: Najas major Allioni, var. paucidentata A. Braun 1864.
Najas major var. ehrenbergii	Najas major Allioni, var. ehrenbergii A. Braun, J. Bot. 2: 275. 1864.
Najas major var. intermedia	Najas major Allioni, var. intermedia (Gorski) A. Braun, J. Bot. 2: 276. 1864.	BASIONYM: Najas intermedia Gorski 1830.
Najas major var. microcarpa	Najas major Allioni, var. microcarpa A. Braun, J. Bot. 2: 276. 1864.		TYPE: CANARY ISLANDS:
Najas major var. multidentata	Najas major Allioni, var. multidentata A. Braun, J. Bot. 2: 275. 1864.		TYPE: FRANCE:
Najas major var. paucidentata	Najas major Allioni, var. paucidentata A. Braun, J. Bot. 2: 276. 1864.		TYPE: FRANCE:
Najas marina subsp. ehrenbergii	Najas marina Linnaeus, subsp. ehrenbergii (A. Braun) L. Triest, Mem. Acad. Roy. Sci. Outre-Mer, Cl. Sci. Nat. Med. Collect. 8vo 21: 24. 1987.	BASIONYM: Najas major Allioni, var. ehrenbergii A. Braun 1864.
Najas marina subsp. intermedia	Najas marina Linnaeus, subsp. intermedia (Gorski) Casper, Feddes Repert. 90: 236. 1979.	BASIONYM: Najas intermedia Gorski 1830.
Najas marina subsp. latior	Najas marina Linnaeus, subsp. latior (F. Mueller ex K. Schmuann) L. Triest, Mém. Acad. Roy. Sci. Outre-Mer, Cl. Sci. Nat. Méd. Collect. 8vo 22: 60. 1988.	BASIONYM: Najas marina Linnaeus, var. latior F. Mueller ex K. Schumann 1894.
Najas marina subsp. microcarpa	Najas marina Linnaeus, subsp. microcarpa (A. Braun) L. Triest, Mém. Acad. Roy. Sci. Outre-Mer, Cl. Sci. Nat. Méd. Collect. 8vo 21: 25. 1987.	BASIONYM: Najas major Allioni, var. microcarpa A. Braun 1864.
Najas marina var. bollei	Najas marina Linnaeus, var. bollei K. Schumann, in Martius, Fl. Bras. 3(3): 726. 1894.		TYPE: CANARY ISLANDS:
Najas marina var. brevifolia	Najas marina Linnaeus, var. brevifolia Rendle, Trans. Linn. Soc. London, Bot. 5: 396. 1899.
Najas marina var. californica	Najas marina Linnaeus, var. californica Rendle, Trans. Linn. Soc. London, Bot. 5: 398. 1899.		TYPE: CALIFORNIA:
Najas marina var. denticulata	Najas marina Linnaeus, var. denticulata Rendle, Trans. Linn. Soc. London, Bot. 5: 438. 1900.
Najas marina var. ehrenbergii	Najas marina Linnaeus, var. ehrenbergii (A. Braun) K. Schumann, in Martius, Fl. Bras. 3(3): 726. 1894.	BASIONYM: Najas major Allioni, var. ehrenbergii A. Braun 1864.
Najas marina var. intermedia	Najas marina Linnaeus, var. intermedia (Gorski) Ascherson, Fl. Brandenburg 1: 670. 1864.	BASIONYM: Najas intermedia Gorski 1830.
Najas marina var. latifolia	Najas marina Linnaeus, var. latifolia (A. Braun) K. Schumann, in Martius, Fl. Bras. 3(3): 725. 1894.	BASIONYM: Najas latifolia A. Braun 1864.
Najas marina var. latior	Najas marina Linnaeus, var. latior F. Mueller ex K. Schumann, in Martius, Fl. Bras. 3(3): 725. 1894.		TYPE: AUSTRALIA: Queensland:
Najas marina var. mexicana	Najas marina Linnaeus, var. mexicana Rendle, Trans. Linn. Soc. London, Bot. 5: 398. 1899.		TYPE: MEXICO: Baja California Sur: Muleje, Gulf of California, Jun 1887, Palmer 20 (holotype: ?; isotypes: NY).
Najas marina var. multidentata	Najas marina Linnaeus, var. multidentata (A. Braun) K. Schumann, in Martius, Fl. Bras. 3(3): 726. 1894.	BASIONYM: Najas major Allioni, var. multidentata A. Braun 1864.
Najas marina var. muricata	Najas marina Linnaeus, var. muricata (Delile) K. Schumann, in Martius, Fl. Bras. 3(3): 725. 1894.	BASIONYM: Najas muricata Delile 1813.
Najas marina var. paucidentata	Najas marina Linnaeus, var. paucidentata (A. Braun) K. Schumann, in Martius, Fl. Bras. 3(3): 726. 1894.	BASIONYM: Najas major Allioni, var. paucidentata A. Braun 1864.
Najas marina var. recurvata	Najas marina Linnaeus, var. recurvata Dudley, Cornell Univ. Sci. Bull. 2: 104. 1886.		TYPE: NEW YORK: Black Lake, Cayuga marshes, 9 Oct 1880, Dudley s.n. (holotype: ?; isotype: NY).
Najas marina var. riedelii	Najas marina Linnaeus, var. riedelii K. Schumann, in Martius, Fl. Bras. 3(3): 725. 1894.		TYPE: BRAZIL: São Paulo: Lagoa de Pertininga {Paratininga}, 1833, Riedel s.n. (lectotype: P). Lectotypified by Lowden, Aquatic Bot. 24: 163. 1986?
Najas microcarpa	Najas microcarpa (A. Braun) Bolle ex Christ, Bot. Jahrb. Syst. 9: 153. 1888.	BASIONYM: Najas major Allioni, var. microcarpa A. Braun 1864.
Najas muricata	Najas muricata Delile, Descr. Egypte, Hist. Nat. 281. 1813 ("1812").

Specimens

Country	State	County	Date	Specimen
USA	Florida	Lee Co.	01 Jan 1976	W. C. Brumbach 8919
USA	Florida	Manatee Co.	28 Oct 1981	A.G. Shuey 2514
USA	Florida	Escambia Co.	16 Jul 1992	P. Hardy s.n.
USA	Florida	Miami-Dade Co.	10 Sep 1966	O. K. Lakela 30204
USA	Florida	Miami-Dade Co.	14 Apr 1948	R. O. Woodbury s.n.
USA	Florida	Sarasota Co.	02 Apr 1981	R. P. Wunderlin 8926
USA	Florida	Lee Co.	22 Nov 1972	K. E. Rogers 8902
USA	Florida	Brevard Co.	02 Aug 1972	A.G. Shuey M1020
USA	Florida	Brevard Co.	02 Aug 1972	A.G. Shuey s.n.
USA	California	San Bernardino Co.	03 Sep 1982	R. F. Thorne 55243
USA	New York	Cayuga Co.	08 Aug 1914	F. P. Metcalf 1516
USA	Arizona	Mohave Co.	12 Nov 1960	R. H. Hevly s.n.
USA	Florida	Hillsborough Co.	14 Jun 2015	S. Dickman 971

Listed Status: Florida

Listed Status:	(Florida):Source - Plants in the Preservation of Native Flora of Florida Act. Chapter 5B-40, Florida Administrative Code. 1998, amended.
Endangered:	A species of plants native to the state that are in imminent danger of extinction within the state, the survival of which is unlikely if the causes of a decline in the number of plants continue, and includes all species determined to be endangered or threatened pursuant to the Federal Endangered Species Act of 1973, as amended.
Threatened:	Listed as Threatened Plants in the Preservation of Native Flora of Florida Act. Defined as species of plants native to the state that are in rapid decline in the number of plants within the state, but which have not so decreased in such number as to cause them to be endangered.

Listed Status: US

Listed Status: US	(U.S.) Source - List of Endangered and Threatened Wildlife and Plants under the provisions of the Endangered Species Act of 1973, as amended.
Endangered:	Any species which is in danger of extinction throughout all or a significant portion of its range.
Threatened:	Any species which is likely to become an endangered species within the foreseeable future throughout all or a significant portion of its range.

WAP: Wetland AssessmentProcedure

Wetland Assessment Procedure (WAP): Source - Southwest Florida Water Management District, Wetland Assessment Procedure Instruction Manual for Isolated Wetlands (March 2005).

AD:	Adaptive Species. Plant species designated as FAC or Upland by DEP, but commonly seen in the transition zone in limited numbers.
D:	Deep Species. Plant species commonly found in the deep zone, and designated either FAC or OBL by DEP.
OD:	Outer Deep Species. Plant species commonly found in the outer deep zone, and designated either FACW or OBL by DEP.
T:	Transition Species. Plant species commonly found in the transition zone, and designated either FACW or OBL by DEP.
U:	Upland Species. Plant species that are not expected to be seen in wetlands.

ANY - WAP critieria is not taken into consideration
Yes - Show results with WAP designations
No - Show results without WAP designations

FISC: Florida Invasive Species Council

FISC:

Source - Florida Invasive Species Council's 2017 List of Florida's Most Invasive Species

Category I - Species that are invading and disrupting native plant communities in Florida. This definition does not rely on the economic severity or geographic range of the problem, but on the documented ecological damage caused.

Category II - Species that have shown a potential to disrupt native plant communities. These species may become ranked as Category I, but have not yet demonstrated disruption of natural Florida communities.

DEP: Department of Environmental Protection

Wetland Status, Department of Environmental Regulation (DEP): Source - Delineation of the Landward Extent of Wetlands and Surface Waters, Chapter 62-340, Florida Administrative Code. 1994.

OBL:	Obligate wetland. Occurs almost always under natural conditions in wetlands.
FACW:	Facultative Wetland. Usually occurs in wetlands, but occasionally found in non-wetlands
FAC:	Facultative. equally likely to occur in wetlands or non-wetlands.

NWPL: National Wetland Plant List

National Wetland Plant List (NWPL): Source - Lichvar, R.W., M. Butterwick, N.C. Melvin, and W.N. Kirchner. 2014. The National Wetland Plant List: 2014 Update of Wetland Ratings. Phytoneuron 2014-41: 1-42.

OBL:	Obligate Wetland. Occurs almost always (estimated probability 99%) under natural conditions in wetlands.
FACW:	Facultative Wetland. Usually occurs in wetlands (estimated probability 67%-99%), but occasionally found in non-wetlands.
FAC:	Facultative. Equally likely to occur in wetlands or non-wetlands (estimated probability 34%-66%).
FACU:	Facultative Upland. Usually occurs in non-wetlands (estimated probability 67%-99%), but occasionally found in wetlands (estimated probability 1%-33%).
UPL:	Obligate Upland. Occurs in wetlands in another region, but occurs almost always (estimated probability 99%) under natural conditions in non-wetlands in the regions specified.

Persisting

Identifying species that appear as waifs or only periodically appear in the flora for a few seasons.

Any - Persisting critieria is not taken into consideration
Yes - Show results with a Persisting value of Y
No - Show results without Persisting value of N

Duration

Annual:	Plants that perform their entire lifecycle within a single growing season. All roots, stems, and leaves die at the end of the growing season. Over wintering seeds allow the next generation to appear.
Biennial:	A plant that is typically vegetative its first year and blooms the following season. Once it has bloomed and set seed, the plant dies. Garlic mustard is an example of a biennial.
Perennial:	These plants live for three or more seasons. Many perennials may not be mature enough to bloom during its first year. Perennial wildflowers re-grow each season from overwinter root material.

Category

Vascular:	Any of various plants that have the vascular tissues xylem and phloem. The vascular plants include all seed-bearing plants (the gymnosperms and angiosperms) and the pteridophytes (including the ferns, lycophytes, and horsetails). Also called tracheophyte.
Bryophyte:	A large group of seedless green plants including the mosses, liverworts, and hornworts. Bryophytes lack the specialized tissues xylem and phloem that circulate water and dissolved nutrients in the vascular plants. Bryophytes generally live on land but are mostly found in moist environments, for they have free-swimming sperm that require water for transport. In contrast to the vascular plants, the gametophyte (haploid) generation of bryophytes constitutes the larger plant form, while the small sporophyte (diploid) generation grows on or within the gametophyte and depends upon it for nutrition.
Lichen:	The mutualistic symbiotic association of a fungus with an alga or a cyanobacterium, or both. The fungal component of a lichen absorbs water and nutrients from the surroundings and provides a suitable environment for the alga or cyanobacterium. These live protected among the dense fungal hyphae and produce carbohydrates for the fungus by photosynthesis. Owing to this partnership, lichens can thrive in harsh environments such as mountaintops and polar regions. The more familiar lichens grow slowly as crusty patches, but lichens are found in a variety of forms, such as the tall, plantlike reindeer moss. The association between the different organisms in a lichen is so close that lichens are routinely referred to as a single organism, and scientists classify lichens using the name of the fungal component.

(Definitions from: American Heritage Science Dictionary)

Growth Habit

Intro paragraph to be provided by New York.

Vine:	DESCRIPTION TO BE PROVIDED BY NEW YORK.
Shrub:	DESCRIPTION TO BE PROVIDED BY NEW YORK.
Tree:	DESCRIPTION TO BE PROVIDED BY NEW YORK.
Et cetera:	DESCRIPTION TO BE PROVIDED BY NEW YORK.

State Rank

This numeric rank provides the relative rarity for each species based on a scale from 1 (very rare) to 5 (common). These ranks carry no legal status.

S1 - Typically 5 or fewer occurrences, very few remaining individuals, acres, or miles of stream, or some factor of its biology making it especially vulnerable in the state.
S2 - Typically 6 to 20 occurrences, few remaining individuals, acres, or miles of stream, or factors demonstrably making it very vulnerable in the state.
S3 - Typically 21 to 100 occurrences, limited acreage, or miles of stream in the state.
S4 - Apparently secure in the state.
S5 - Demonstrably secure in the state.
SE - State exotic or non-native.
SH - Historically known from the state, but not seen in the past 15 years.
SNA - Species for which a rank is not applicable. This is mainly those species which are now excluded from flora for various reasons.
SNR - Not yet ranked.
SX - Apparently extirpated from the state.

Global Rank

Each species' global rank is determined by NatureServe. These ranks carry no legal weight. The global rank reflects the species worldwide rarity.

G1 - Critically imperiled globally because of extreme rarity (5 or fewer occurrences), or very few remaining acres, or miles of stream) or especially vulnerable to extinction because of some factor of its biology.
G2 - Imperiled globally because of rarity (6 - 20 occurrences, or few remaining acres, or miles of stream) or very vulnerable to extinction throughout its range because of other factors.
G3 - Imperiled globally because of rarity (6 - 20 occurrences, or few remaining acres, or miles of stream) or very vulnerable to extinction throughout its range because of other factors.
G4 - Imperiled globally because of rarity (6 - 20 occurrences, or few remaining acres, or miles of stream) or very vulnerable to extinction throughout its range because of other factors.
G5 - Demonstrably secure globally, though it may be quite rare in parts of its range, especially at the periphery.
GH - Historically known, with the expectation that it might be rediscovered.
GNA - Species for which a rank is not applicable. NatureServe does not typically rank hybrid species.
GNR - Not yet ranked.
GX - Species believed to be extinct.

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Associated Ecological Communities

View a List of All Ecological Communities

For more information, contact: Richard Wunderlin or Bruce Hansen, Alan Franck, or Karla Alvarado

Web Development: USF Water Institute

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Citation Information:

Wunderlin, R. P., B. F. Hansen, A. R. Franck, and F. B. Essig. 2024. Atlas of Florida Plants (http://florida.plantatlas.usf.edu/). [S. M. Landry and K. N. Campbell (application development), USF Water Institute.] Institute for Systematic Botany, University of South Florida, Tampa.

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Definition: Vouchered Specimen

A voucher specimen is a pressed and thoroughly dried plant sample deposited in a herbarium, and is intended to be a permanent record supporting research purposes. A voucher may be a record of a plant's occurrence in a particular area, or a specific example of a plant used in a scientific investigation.

Proper vouchers display all the necessary attributes for complete identification of the plant, and are to be accompanied by accurate locality, habitat, collection time, and collector data.

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