Euphorbia spathulata

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Family:	EUPHORBIACEAE
Species:	Euphorbia spathulata Lam.
Common Name:	WARTY SPURGE; PRAIRIE SPURGE
Status:	Native, FACU (NWPL)
Specimen:	View details of USF Herbarium specimens

** Not applicable or data not available.

Classification

Division
Class
Sub Class
Order	MALPIGHIALES
Family	EUPHORBIACEAE
Genus	Euphorbia
Species	Euphorbia spathulata Lam. - WARTY SPURGE; PRAIRIE SPURGE

Citation

Citation	EUPHORBIA SPATHULATA Lamarck, Encycl. 2: 428. 1788.
Basionym:	**
Type:	URUGUAY: Near Montevideo, s.d., Commerson s.n. (holotype: P; isotype: MPU).

** Not applicable or data not available.

Source

County	Herbaria	Literature Citation (If Applicable)
Escambia	FSU

Synonyms

– Denotes synonyms that are applicable to the state. Show these synonyms only

Synonym	Full Citation	Basionym	Type
Tithymalus spathulatus	Tithymalus spathulatus (Lamarck) W. A. Weber, Phytologia 58: 384. 1985.	BASIONYM: Euphorbia spathulata Lamarck 1788.
Xamesike deltifolia	Xamesike deltifolia Rafinesque, Autik. Bot. 98. 1840.		TYPE: FLORIDA:
Euphorbia arkansana	Euphorbia arkansana Engelmann & A. Gray, Boston J. Nat. Hist. 5: 261. 1845.		TYPE: TEXAS: Harris Co.: Comal Spring, near Houston, Jun 1845, Lindheimer s.n. [Fl. Tex. Exsicc. 302] (lectotype: MO; isolectotypes: GH, LE, NY). Lectotypified by Geltman et al., J. Bot. Res. Inst. Texas 5: 148. 2011.
Euphorbia arkansana var. atrosemina	Euphorbia arkansana Engelmann & A. Gray, var. atrosemina Norton, N. Amer. Euphorbia 21, pl. 20. 1899.		TYPE: ARIZONA: Foothills of the Santa Rita Mountains, 19 May 1894, Pringle s.n. (lectotype: US; isolectotype: GH). Lectotypified by Geltman et al., J. Bot. Res. Inst. Texas 5: 149. 2011.
Euphorbia arkansana var. coloradensis	Euphorbia arkansana Engelmann & A. Gray, var. coloradensis Norton, N. Amer. Euphorbia 21, pl. 19. 1899.		TYPE: COLORADO: Clear Creek Co.: Clear Creek Canyon, 20 Jun 1878, Jones 239 (lectotype: MO). Lectotypified by Geltman et al., J. Bot. Res. Inst. Texas 5: 149. 2011.
Euphorbia arkansana var. missouriensis	Euphorbia arkansana Engelmann & A. Gray, var. missouriensis Norton, N. Amer. Euphorbia 19. 1899.		TYPE: KANSAS: Riley Co.: Manhattan, 11 Jun 1897, Norton s.n. (lectotype: MO). Lectotypified by Geltman et al., J. Bot. Res. Inst. Texas 5: 149. 2011.
Euphorbia dictyosperma	Euphorbia dictyosperma Fischer & C. A. Meyer, Index Sem. Hort. Petrop. 2: 37. 1836.		TYPE: CALIFORNIA: Cult. in Russia, 1835, Meyer s.n. (lectotype: LE). Lectotypified by Geltman et al., J. Bot. Res. Inst. Texas 5: 148. 2011.
Euphorbia dictyosperma var. mexicana	Euphorbia dictyosperma Fischer & C. A. Meyer, var. mexicana Engelmann, in Emory, Rep. U.S. Mex. Bound. 2(1): 191. 1858.		TYPE: MEXICO: Durango: Bolsón de Mapimí, Valle de Nazas, 15 Apr 1847, Gregg 456 (lectotype: MO). Lectotypified by S. J. Wolf, Ann. Missouri Bot. Gard. 75: 1626. 1988.
Euphorbia dictyosperma var. multicaulis	Euphorbia dictyosperma Fischer & C. A Meyer, var. multicaulis J. M. Coulter, Contr. U.S. Natl. Herb. 2: 393. 1894.	BASIONYM: Euphorbia multicaulis Engelmann 1858, non Thuillier 1799.
Euphorbia mexicana	Euphorbia mexicana (Engelmann) Norton, N. Amer. Euphorbia 21. 1899.	BASIONYM: Euphorbia dictyosperma Fischer & C. A. Meyer, var. mexicana Engelmann 1858.
Euphorbia missouriensis	Euphorbia missouriensis (Norton) Small, in Britton, Man. Fl. N. States 595. 1901.	BASIONYM: Euphorbia arkansana Engelmann & A. Gray, var. missouriensis Norton 1899.
Euphorbia multicaulis	Euphorbia multicaulis Engelmann, in Emory, Rep. U.S. Mex. Bound. 2(1): 191. 1858, non Thuillier 1799.		TYPE: MEXICO: Sonora: Las Playas, May 1851, Thurber 381 (holotype: MO).
Euphorbia obtusata	Euphorbia obtusata Pursh, Fl. Amer. Sept. 606. 1814.		TYPE: VIRGINIA: August Co.: Staunton, 1806, Pursh 116 (lectotype: PH 1021023, rightmost fragment). Lectotypified by Geltman et al., J. Bot. Res. Inst. Texas 5: 148. 2011.
Euphorbia pseudopeplus	Euphorbia pseudopeplus Spegazzini, Anales Mus. Nac. Buenos Aires 7: 164. 1902.		TYPE: ARGENTINA: Río Negro: Dpto. Adolfo Alsina, near Carmen de Patagones, Feb 1898, Spegazzini s.n. [Herb. Spegazzini no. 12560] (lectotype: LP). Lectotypified by Geltman et al., J. Bot. Res. Inst. Texas 5: 149. 2011.
Euphorbia spathulata var. mexicana	Euphorbia spathulata Lamarck, var. mexicana (Engelmann) Oudejans, Collect. Bot. (Barcelona) 21: 188. 1993 ("1992").	BASIONYM: Euphorbia dictyosperma Fischer & C. A. Meyer, var. mexicana Engelmann 1858.
Galarhoeus arkansanus	Galarhoeus arkansanus (Engelmann & A. Gray) Small ex Rydberg, Fl. Plains N. Amer. 520. 1932.	BASIONYM: Euphorbia arkansana Engelmann & A. Gray 1845.
Galarhoeus missouriensis	Galarhoeus missouriensis (Norton) Rydberg, Fl. Plains N. Amer. 521. 1932.	BASIONYM: Euphorbia arkansana Engelmann & A. Gray, var. missouriensis Norton 1899.
Galarhoeus obtusatus	Galarhoeus obtusatus (Pursh) Small ex Rydberg, Fl. Plains N. Amer. 520. 1932.	BASIONYM: Euphorbia obtusata Pursh 1814.
Keraselma spatulatus	Keraselma spathulatus Rafinesque, Autik. Bot. 93. 1840.
Tithymalus arkansanus	Tithymalus arkansanus (Engelmann & A. Gray) Klotzsch & Garcke, in Klotzsch, Abh. Königl. Akad. Wiss. Berlin 1859(1): 66. 1860.	BASIONYM: Euphorbia arkansana Engelmann & A. Gray 1845.
Tithymalus arkansanus var. coloradensis	Tithymalus arkansanus (Engelmann & A. Gray) Klotzsch & Garcke, var. coloradensis (Norton) Rydberg, Bull. Torrey Bot. Club 33: 145. 1906.	BASIONYM: Euphorbia arkansana Engelmann & A. Gray, var. coloradensis Norton 1899.
Tithymalus dictyospermus	Tithymalus dictyospermus (Fischer & C. A. Meyer) A. Heller, Muhlenbergia 1: 56. 1904.	BASIONYM: Euphorbia dictyosperma Fischer & C. A. Meyer 1836.
Tithymalus mexicanus	Tithymalus mexicanus (Engelmann) Wooton & Standley, Contr. U.S. Natl. Herb. 16: 145. 1913.	BASIONYM: Euphorbia dictyosperma Fischer & C. A. Meyer, var. mexicana Engelmann 1858.
Tithymalus missouriensis	Tithymalus missouriensis (Norton) Small, Fl. S.E. U.S. 721, 1334. 1903.	BASIONYM: Euphorbia arkansana Engelmann & A. Gray, var. missouriensis Norton 1899.
Tithymalus obtusatus	Tithymalus obtusatus (Pursh) Klotzsch & Garcke, in Klotzsch, Abh. Königl. Akad. Wiss. Berlin 1859(1): 69. 1860.	BASIONYM: Euphorbia obtusata Pursh 1814.
Tithymalus pseudopeplus	Tithymalus pseudopeplus (Spegazzini) Soják, Cas Nár. Mus., Odd. Prír. 148: 199. 1980 ("1979").	BASIONYM: Euphorbia pseudopeplus Spegazzini 1902.

Specimens

Country	State	County	Date	Specimen	Notes
USA	South Carolina	Pickens Co.	15 May 1994	S. R. Hill 25575	CULTIVATED; from Horry County.
USA	Louisiana	St. Martin Par.	18 May 1964	J. W. Thieret 17375
USA	Louisiana	East Baton Rouge Par.	17 Mar 1976	M. Schmitt 40
USA	North Carolina	Orange Co.	29 May 1959	R. L. Wilbur 6225
USA	North Carolina	Perquimans co.	09 May 1958	H. E. Ahles 39926
USA	Mississippi	Clay Co.	05 May 1955	J. D. Ray, Jr. 4411
USA	Alabama	Colbert Co.	17 Apr 1955	G. R. Cooley 3486
USA	Oklahoma	Murray Co.	05 Apr 1975	R. J. Tyrl 958
USA	Texas	Garza Co.	16 Apr 1963	V. de Vlaming 88
USA	Texas	Brown Co.	08 Apr 1968	J. W. Stanford 1809

Listed Status: Florida

Listed Status:	(Florida):Source - Plants in the Preservation of Native Flora of Florida Act. Chapter 5B-40, Florida Administrative Code. 1998, amended.
Endangered:	A species of plants native to the state that are in imminent danger of extinction within the state, the survival of which is unlikely if the causes of a decline in the number of plants continue, and includes all species determined to be endangered or threatened pursuant to the Federal Endangered Species Act of 1973, as amended.
Threatened:	Listed as Threatened Plants in the Preservation of Native Flora of Florida Act. Defined as species of plants native to the state that are in rapid decline in the number of plants within the state, but which have not so decreased in such number as to cause them to be endangered.

Listed Status: US

Listed Status: US	(U.S.) Source - List of Endangered and Threatened Wildlife and Plants under the provisions of the Endangered Species Act of 1973, as amended.
Endangered:	Any species which is in danger of extinction throughout all or a significant portion of its range.
Threatened:	Any species which is likely to become an endangered species within the foreseeable future throughout all or a significant portion of its range.

WAP: Wetland AssessmentProcedure

Wetland Assessment Procedure (WAP): Source - Southwest Florida Water Management District, Wetland Assessment Procedure Instruction Manual for Isolated Wetlands (March 2005).

AD:	Adaptive Species. Plant species designated as FAC or Upland by DEP, but commonly seen in the transition zone in limited numbers.
D:	Deep Species. Plant species commonly found in the deep zone, and designated either FAC or OBL by DEP.
OD:	Outer Deep Species. Plant species commonly found in the outer deep zone, and designated either FACW or OBL by DEP.
T:	Transition Species. Plant species commonly found in the transition zone, and designated either FACW or OBL by DEP.
U:	Upland Species. Plant species that are not expected to be seen in wetlands.

ANY - WAP critieria is not taken into consideration
Yes - Show results with WAP designations
No - Show results without WAP designations

FISC: Florida Invasive Species Council

FISC:

Source - Florida Invasive Species Council's 2017 List of Florida's Most Invasive Species

Category I - Species that are invading and disrupting native plant communities in Florida. This definition does not rely on the economic severity or geographic range of the problem, but on the documented ecological damage caused.

Category II - Species that have shown a potential to disrupt native plant communities. These species may become ranked as Category I, but have not yet demonstrated disruption of natural Florida communities.

DEP: Department of Environmental Protection

Wetland Status, Department of Environmental Regulation (DEP): Source - Delineation of the Landward Extent of Wetlands and Surface Waters, Chapter 62-340, Florida Administrative Code. 1994.

OBL:	Obligate wetland. Occurs almost always under natural conditions in wetlands.
FACW:	Facultative Wetland. Usually occurs in wetlands, but occasionally found in non-wetlands
FAC:	Facultative. equally likely to occur in wetlands or non-wetlands.

NWPL: National Wetland Plant List

National Wetland Plant List (NWPL): Source - Lichvar, R.W., M. Butterwick, N.C. Melvin, and W.N. Kirchner. 2014. The National Wetland Plant List: 2014 Update of Wetland Ratings. Phytoneuron 2014-41: 1-42.

OBL:	Obligate Wetland. Occurs almost always (estimated probability 99%) under natural conditions in wetlands.
FACW:	Facultative Wetland. Usually occurs in wetlands (estimated probability 67%-99%), but occasionally found in non-wetlands.
FAC:	Facultative. Equally likely to occur in wetlands or non-wetlands (estimated probability 34%-66%).
FACU:	Facultative Upland. Usually occurs in non-wetlands (estimated probability 67%-99%), but occasionally found in wetlands (estimated probability 1%-33%).
UPL:	Obligate Upland. Occurs in wetlands in another region, but occurs almost always (estimated probability 99%) under natural conditions in non-wetlands in the regions specified.

Persisting

Identifying species that appear as waifs or only periodically appear in the flora for a few seasons.

Any - Persisting critieria is not taken into consideration
Yes - Show results with a Persisting value of Y
No - Show results without Persisting value of N

Duration

Annual:	Plants that perform their entire lifecycle within a single growing season. All roots, stems, and leaves die at the end of the growing season. Over wintering seeds allow the next generation to appear.
Biennial:	A plant that is typically vegetative its first year and blooms the following season. Once it has bloomed and set seed, the plant dies. Garlic mustard is an example of a biennial.
Perennial:	These plants live for three or more seasons. Many perennials may not be mature enough to bloom during its first year. Perennial wildflowers re-grow each season from overwinter root material.

Category

Vascular:	Any of various plants that have the vascular tissues xylem and phloem. The vascular plants include all seed-bearing plants (the gymnosperms and angiosperms) and the pteridophytes (including the ferns, lycophytes, and horsetails). Also called tracheophyte.
Bryophyte:	A large group of seedless green plants including the mosses, liverworts, and hornworts. Bryophytes lack the specialized tissues xylem and phloem that circulate water and dissolved nutrients in the vascular plants. Bryophytes generally live on land but are mostly found in moist environments, for they have free-swimming sperm that require water for transport. In contrast to the vascular plants, the gametophyte (haploid) generation of bryophytes constitutes the larger plant form, while the small sporophyte (diploid) generation grows on or within the gametophyte and depends upon it for nutrition.
Lichen:	The mutualistic symbiotic association of a fungus with an alga or a cyanobacterium, or both. The fungal component of a lichen absorbs water and nutrients from the surroundings and provides a suitable environment for the alga or cyanobacterium. These live protected among the dense fungal hyphae and produce carbohydrates for the fungus by photosynthesis. Owing to this partnership, lichens can thrive in harsh environments such as mountaintops and polar regions. The more familiar lichens grow slowly as crusty patches, but lichens are found in a variety of forms, such as the tall, plantlike reindeer moss. The association between the different organisms in a lichen is so close that lichens are routinely referred to as a single organism, and scientists classify lichens using the name of the fungal component.

(Definitions from: American Heritage Science Dictionary)

Growth Habit

Intro paragraph to be provided by New York.

Vine:	DESCRIPTION TO BE PROVIDED BY NEW YORK.
Shrub:	DESCRIPTION TO BE PROVIDED BY NEW YORK.
Tree:	DESCRIPTION TO BE PROVIDED BY NEW YORK.
Et cetera:	DESCRIPTION TO BE PROVIDED BY NEW YORK.

State Rank

This numeric rank provides the relative rarity for each species based on a scale from 1 (very rare) to 5 (common). These ranks carry no legal status.

S1 - Typically 5 or fewer occurrences, very few remaining individuals, acres, or miles of stream, or some factor of its biology making it especially vulnerable in the state.
S2 - Typically 6 to 20 occurrences, few remaining individuals, acres, or miles of stream, or factors demonstrably making it very vulnerable in the state.
S3 - Typically 21 to 100 occurrences, limited acreage, or miles of stream in the state.
S4 - Apparently secure in the state.
S5 - Demonstrably secure in the state.
SE - State exotic or non-native.
SH - Historically known from the state, but not seen in the past 15 years.
SNA - Species for which a rank is not applicable. This is mainly those species which are now excluded from flora for various reasons.
SNR - Not yet ranked.
SX - Apparently extirpated from the state.

Global Rank

Each species' global rank is determined by NatureServe. These ranks carry no legal weight. The global rank reflects the species worldwide rarity.

G1 - Critically imperiled globally because of extreme rarity (5 or fewer occurrences), or very few remaining acres, or miles of stream) or especially vulnerable to extinction because of some factor of its biology.
G2 - Imperiled globally because of rarity (6 - 20 occurrences, or few remaining acres, or miles of stream) or very vulnerable to extinction throughout its range because of other factors.
G3 - Imperiled globally because of rarity (6 - 20 occurrences, or few remaining acres, or miles of stream) or very vulnerable to extinction throughout its range because of other factors.
G4 - Imperiled globally because of rarity (6 - 20 occurrences, or few remaining acres, or miles of stream) or very vulnerable to extinction throughout its range because of other factors.
G5 - Demonstrably secure globally, though it may be quite rare in parts of its range, especially at the periphery.
GH - Historically known, with the expectation that it might be rediscovered.
GNA - Species for which a rank is not applicable. NatureServe does not typically rank hybrid species.
GNR - Not yet ranked.
GX - Species believed to be extinct.

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Filter by County - Counties to Exclude - ANY versus ALL

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Associated Ecological Communities

View a List of All Ecological Communities

For more information, contact: Richard Wunderlin or Bruce Hansen, Alan Franck, or Karla Alvarado

Web Development: USF Water Institute

A member of the University of South Florida family of PlantAtlas.org websites

Citation Information:

Wunderlin, R. P., B. F. Hansen, A. R. Franck, and F. B. Essig. 2025. Atlas of Florida Plants (http://florida.plantatlas.usf.edu/). [S. M. Landry and K. N. Campbell (application development), USF Water Institute.] Institute for Systematic Botany, University of South Florida, Tampa.

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Definition: Vouchered Specimen

A voucher specimen is a pressed and thoroughly dried plant sample deposited in a herbarium, and is intended to be a permanent record supporting research purposes. A voucher may be a record of a plant's occurrence in a particular area, or a specific example of a plant used in a scientific investigation.

Proper vouchers display all the necessary attributes for complete identification of the plant, and are to be accompanied by accurate locality, habitat, collection time, and collector data.

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